There were two overall runners up and winners from five categories: Community, population and macroecology behavioural ecology and physiology, conservation ecology and biodiversity, landscape ecology and ecosystems and the editor's Pick. It defeated 139 other images to the title and judges said it does a 'fantastic job of showcasing the striking beauty of a rather enigmatic species'. It was captured Southern Andes of Ecuador two hours after the butterfly emerged from its chrysalis (right) - the third and final life stage before adulthood where the insect is a pupa.
